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In cladistics, as theorised by Willi Hennig (cf. Schmitt 2013), a character or a character state may be plesiomorphic (ancestral or primitive) or apomorphic (derived). The polarisation of characters (the determination of the direction of character change), which is at the core of the phylogenetic method, is comparable to the concept of "error of copying" in the Lachmann's method. An apomorphic character state is equivalent to a secondary reading.